"\u0420\u043E\u0301\u0431\u0435\u0440\u0442 \u0412\u0456\u0301\u043B\u044C\u044F\u043C \u0413\u043E\u0301\u043B\u043B\u0456 (\u0430\u043D\u0433\u043B. Robert William Holley; 28 \u0441\u0456\u0447\u043D\u044F 1922, \u0423\u0440\u0431\u0430\u043D\u0430, \u0406\u043B\u043B\u0456\u043D\u043E\u0439\u0441 \u2014 11 \u043B\u044E\u0442\u043E\u0433\u043E 1993, \u041B\u043E\u0441-\u0413\u0430\u0442\u043E\u0441, \u041A\u0430\u043B\u0456\u0444\u043E\u0440\u043D\u0456\u044F, \u0421\u0428\u0410) \u2014 \u0430\u043C\u0435\u0440\u0438\u043A\u0430\u043D\u0441\u044C\u043A\u0438\u0439 \u0431\u0456\u043E\u0445\u0456\u043C\u0456\u043A, \u043B\u0430\u0432\u0440\u0435\u0430\u0442 \u041D\u043E\u0431\u0435\u043B\u0456\u0432\u0441\u044C\u043A\u043E\u0457 \u043F\u0440\u0435\u043C\u0456\u0457 \u0437 \u043C\u0435\u0434\u0438\u0446\u0438\u043D\u0438 \u0442\u0430 \u0444\u0456\u0437\u0456\u043E\u043B\u043E\u0433\u0456\u0457 1968 \u0440\u043E\u043A\u0443 (\u0441\u043F\u0456\u043B\u044C\u043D\u043E \u0437 \u0413\u0430\u0440\u043E\u043C \u0425\u043E\u0440\u0430\u043D\u0430 \u0456 \u041C\u0430\u0440\u0448\u0430\u043B\u043B\u043E\u043C \u041D\u0456\u0440\u0435\u043D\u0431\u0435\u0440\u0433\u043E\u043C) \u00AB\u0437\u0430 \u0440\u043E\u0437\u0448\u0438\u0444\u0440\u043E\u0432\u043A\u0443 \u0433\u0435\u043D\u0435\u0442\u0438\u0447\u043D\u043E\u0433\u043E \u043A\u043E\u0434\u0443 \u0456 \u0439\u043E\u0433\u043E \u0440\u043E\u043B\u0456 \u0432 \u0441\u0438\u043D\u0442\u0435\u0437\u0456 \u0431\u0456\u043B\u043A\u0456\u0432\u00BB, \u0437\u043E\u043A\u0440\u0435\u043C\u0430 \u0432\u043D\u0435\u0441\u043A\u043E\u043C \u0413\u043E\u043B\u043B\u0456 \u0431\u0443\u0432 \u043E\u043F\u0438\u0441 \u043F\u043E\u0441\u043B\u0456\u0434\u043E\u0432\u043D\u043E\u0441\u0442\u0456 \u0439 \u0441\u0442\u0440\u0443\u043A\u0442\u0443\u0440\u0438 \u0442\u0420\u041D\u041A \u0430\u043B\u0430\u043D\u0456\u043D\u0443, \u043F\u0435\u0440\u0448\u043E\u0457 \u043C\u043E\u043B\u0435\u043A\u0443\u043B\u0438 \u0420\u041D\u041A, \u0434\u043B\u044F \u044F\u043A\u043E\u0457 \u0431\u0443\u043B\u0430 \u0432\u0441\u0442\u0430\u043D\u043E\u0432\u043B\u0435\u043D\u0430 \u0441\u0442\u0440\u0443\u043A\u0442\u0443\u0440\u0430."@uk . . . . . . . . . "Robert William Holley, f\u00F6dd 28 januari 1922 i Urbana, Illinois, USA, d\u00F6d 11 februari 1993 i Los Gatos, Kalifornien, var en amerikansk biokemist. \u00C5r 1968 erh\u00F6ll han Nobelpriset i fysiologi eller medicin tillsammans med Har Gobind Khorana och Marshall W. Nirenberg f\u00F6r deras tolkning av den genetiska koden och ha beskrivit strukturen f\u00F6r alanin\u00F6verf\u00F6rings-RNA, som l\u00E4nkar DNA och proteinsyntes. Vidare tilldelades han 1965 Albert Lasker Basic Medical Research Award."@sv . . . "\u0420\u043E\u0301\u0431\u0435\u0440\u0442 \u0423\u0438\u0301\u043B\u044C\u044F\u043C \u0425\u043E\u0301\u043B\u043B\u0438 (\u0430\u043D\u0433\u043B. Robert William Holley; 28 \u044F\u043D\u0432\u0430\u0440\u044F 1922, \u042D\u0440\u0431\u0430\u043D\u0430, \u0418\u043B\u043B\u0438\u043D\u043E\u0439\u0441 \u2014 11 \u0444\u0435\u0432\u0440\u0430\u043B\u044F 1993, \u041B\u043E\u0441-\u0413\u0430\u0442\u043E\u0441, \u041A\u0430\u043B\u0438\u0444\u043E\u0440\u043D\u0438\u044F) \u2014 \u0430\u043C\u0435\u0440\u0438\u043A\u0430\u043D\u0441\u043A\u0438\u0439 \u0431\u0438\u043E\u0445\u0438\u043C\u0438\u043A, \u043B\u0430\u0443\u0440\u0435\u0430\u0442 \u041D\u043E\u0431\u0435\u043B\u0435\u0432\u0441\u043A\u043E\u0439 \u043F\u0440\u0435\u043C\u0438\u0438 \u043F\u043E \u0444\u0438\u0437\u0438\u043E\u043B\u043E\u0433\u0438\u0438 \u0438 \u043C\u0435\u0434\u0438\u0446\u0438\u043D\u0435 \u0432 1968 \u0433\u043E\u0434\u0443 (\u0441\u043E\u0432\u043C\u0435\u0441\u0442\u043D\u043E \u0441 \u0425\u0430\u0440\u043E\u043C \u041A\u043E\u0440\u0430\u043D\u0430 \u0438 \u041C\u0430\u0440\u0448\u0430\u043B\u043B\u043E\u043C \u041D\u0438\u0440\u0435\u043D\u0431\u0435\u0440\u0433\u043E\u043C) \u00AB\u0437\u0430 \u0440\u0430\u0441\u0448\u0438\u0444\u0440\u043E\u0432\u043A\u0443 \u0433\u0435\u043D\u0435\u0442\u0438\u0447\u0435\u0441\u043A\u043E\u0433\u043E \u043A\u043E\u0434\u0430 \u0438 \u0435\u0433\u043E \u0440\u043E\u043B\u0438 \u0432 \u0441\u0438\u043D\u0442\u0435\u0437\u0435 \u0431\u0435\u043B\u043A\u043E\u0432\u00BB. \u0427\u043B\u0435\u043D \u041D\u0430\u0446\u0438\u043E\u043D\u0430\u043B\u044C\u043D\u043E\u0439 \u0430\u043A\u0430\u0434\u0435\u043C\u0438\u0438 \u043D\u0430\u0443\u043A \u0421\u0428\u0410 (1968)."@ru . "Robert W. Holley"@de . . . . "\u0631\u0648\u0628\u0631\u062A \u0648\u0644\u064A\u0627\u0645 \u0647\u0648\u0644\u064A (\u0628\u0627\u0644\u0625\u0646\u062C\u0644\u064A\u0632\u064A\u0629: Robert W. Holley)\u200F \u0643\u0627\u0646 \u0639\u0627\u0644\u0645 \u0643\u064A\u0645\u064A\u0627\u0621 \u062D\u064A\u0648\u064A\u0629 \u0623\u0645\u0631\u064A\u0643\u064A (28 \u064A\u0646\u0627\u064A\u0631 1922 - 11 \u0641\u0628\u0631\u0627\u064A\u0631 1993)\u060C \u062A\u062D\u0635\u0644 \u0639\u0644\u0649 \u062C\u0627\u0626\u0632\u0629 \u0646\u0648\u0628\u0644 \u0641\u064A \u0627\u0644\u0637\u0628 \u0644\u0639\u0627\u0645 1968 \u0645\u0634\u0627\u0631\u0643\u0629 \u0645\u0639 \u0647\u0627\u0631 \u063A\u0648\u0628\u0646\u062F \u062E\u0648\u0631\u0627\u0646\u0627 \u0648\u0645\u0627\u0631\u0634\u0627\u0644 \u0646\u064A\u0631\u0646\u0628\u0631\u063A \u0648\u0630\u0644\u0643 \u0644\u0623\u0628\u062D\u0627\u062B\u0647\u0645 \u0627\u0644\u062A\u064A \u0623\u0644\u0642\u062A \u0627\u0644\u0636\u0648\u0621 \u0639\u0644\u0649 \u0627\u0644\u0645\u0643\u0648\u0646\u0627\u062A \u0627\u0644\u0648\u0631\u0627\u062B\u064A\u0629 \u0627\u0644\u0645\u0633\u0626\u0648\u0644\u0629 \u0639\u0646 \u062A\u0646\u0638\u064A\u0645 \u0639\u0645\u0644\u064A\u0629 \u062A\u062E\u0644\u064A\u0642 \u0627\u0644\u0628\u0631\u0648\u062A\u064A\u0646\u0627\u062A \u0641\u064A \u0646\u0648\u0627\u0629 \u0627\u0644\u062E\u0644\u064A\u0629."@ar . . . . . . . . "Robert William Holley, f\u00F6dd 28 januari 1922 i Urbana, Illinois, USA, d\u00F6d 11 februari 1993 i Los Gatos, Kalifornien, var en amerikansk biokemist. \u00C5r 1968 erh\u00F6ll han Nobelpriset i fysiologi eller medicin tillsammans med Har Gobind Khorana och Marshall W. Nirenberg f\u00F6r deras tolkning av den genetiska koden och ha beskrivit strukturen f\u00F6r alanin\u00F6verf\u00F6rings-RNA, som l\u00E4nkar DNA och proteinsyntes. Vidare tilldelades han 1965 Albert Lasker Basic Medical Research Award."@sv . "Robert William HOLLEY (28a de Januaro, 1922 \u2013 11a de Februaro, 1993) estis usona biokemiisto. Li kunhavis la Nobel-premion pri fiziologio a\u016D medicino en 1968 (kun Har Gobind Khorana kaj Marshall Warren Nirenberg) \u0109ar ili priskribis la strukturon de alanintransiga RNA, ligante DNA kaj proteina sintezo. Holley naski\u011Dis en Urbana, Ilinojso, kaj gradi\u011Dis el la Urbana altlernejo en 1938. Li studs kemion en la , kaj gradi\u011Dis en 1942 kaj komencis siajn FD studojn en organika kemio \u0109e la Universitato Cornell. Dum la Dua Mondmilito Holley pasigis du jarojn laborante kun la Profesoro Vincent du Vigneaud en la Medicina Kolegio de la Universitato Cornell, kie li eniris en la unua kemia sintezo de penicilino. Holley kompletigis siajn doktorigajn studojn en 1947. Post siaj gradigaj studoj Holley restis asocia kun Cornell. Li i\u011Dis helpoprofesoro de organika kemio en 1948, kaj estis nomumita Profesoro de Biokemio en 1962. Li komencis sian esploradon pri RNA post pasigi sabatan jaron (1955\u20131956) studante kun James F. Bonner en la Kalifornia Instituto de Teknologio. La strukturo de la alanina t-RNA estis kompletigita en 1964, kaj tio estis \u015Dlosila malkovro por klarigi la sintezon de proteinoj el mesa\u011Da RNA. \u011Ci estis anka\u016D la unua nukleotida sekvenco de RNA iam determinita. Holley ricevis la Nobel-premion pri fiziologio a\u016D medicino en 1968 pro tiu malkovro, kaj anka\u016D Har Gobind Khorana kaj Marshall W. Nirenberg ricevis la saman premion tiun jaron pro kontribuoj al la kompreno de la proteinsintezoj."@eo . "Robert W. Holley"@cs . . "\u7F85\u4F2F\u7279\u00B7\u5A01\u5EC9\u00B7\u970D\u5229\uFF08\u82F1\u8A9E\uFF1ADr Robert William Holley\uFF0C1922\u5E741\u670828\u65E5\uFF0D1993\u5E742\u670811\u65E5\uFF09\uFF0C\u7F8E\u570B\u751F\u7269\u5316\u5B78\u5BB6\uFF0C\u51FA\u751F\u65BC\u4F0A\u5229\u8AFE\u5DDE\u5384\u5DF4\u7D0D\u3002\u66FE\u7372\u5F971968\u5E74\u8AFE\u8C9D\u723E\u751F\u7406\u5B78\u6216\u91AB\u5B78\u734E\uFF0C\u539F\u56E0\u662F\u95E1\u660E\u4E86\u9023\u7D50\u4E19\u6C28\u9178\u8207DNA\u7684tRNA\u3002\u5171\u540C\u7372\u734E\u4EBA\u9084\u6709\u54C8\u723E\u00B7\u845B\u8CD3\u00B7\u79D1\u62C9\u7D0D\uFF08Har Gobind Khorana\uFF09\u53CA\u99AC\u6B47\u723E\u00B7\u6C83\u502B\u00B7\u5C3C\u502B\u4F2F\u683C\uFF08Marshall Warren Nirenberg\uFF09\u3002"@zh . . . . . "Robert W. Holley"@en . . "1993-02-11"^^ . "Robert Holley"@pt . . . . . "Robert William Holley (Urbana, Illinois, EUA 1922 - Los Gatos 1993) fou un bioqu\u00EDmic i professor universitari nord-americ\u00E0 guardonat amb el Premi Nobel de Medicina o Fisiologia l'any 1968."@ca . . . . . . "Robert William Holley (28. ledna 1922 \u2013 11. \u00FAnora 1993) byl americk\u00FD biochemik a chemik, nositel Nobelovy ceny za fyziologii a l\u00E9ka\u0159stv\u00ED za rok 1968. Spolu s n\u00EDm ji z\u00EDskali Har Gobind Khorana a Marshall Warren Nirenberg a cena byla ud\u011Blena za vysv\u011Btlen\u00ED toho, jak sekvence nukleotid\u016F v nukleov\u00FDch kyselin\u00E1ch \u0159\u00EDd\u00ED synt\u00E9zu protein\u016F v bu\u0148k\u00E1ch. V\u011Bt\u0161inu sv\u00E9 v\u011Bdeck\u00E9 kari\u00E9ry str\u00E1vil Robert W. Holley na Cornellov\u011B univerzit\u011B, kde i studoval."@cs . . . . "Robert William Holley (Urbana (Illinois) 28 januari 1922 \u2013 Los Gatos 11 februari 1993) was een Amerikaans biochemicus en Nobelprijswinnaar. In 1968 won hij samen met Har Gobind Khorana en Marshall Nirenberg de Nobelprijs voor de Fysiologie of Geneeskunde voor het beschrijven van de structuur van alanine transfer RNA, verbindend DNA en prote\u00EFnesynthese."@nl . . . . "Robert William Holley"@pl . . . . . . . . . . "Robert William Holley (ur. 28 stycznia 1922 w Urbana, zm. 11 lutego 1993 w Los Gatos) \u2013 ameryka\u0144ski biochemik, laureat Nagrody Nobla z dziedziny fizjologii lub medycyny w roku 1968 za wyja\u015Bnienie istoty kodu genetycznego i ustalenie struktury tRNA."@pl . . . "1508474"^^ . "1922-01-28"^^ . . . "\uB85C\uBC84\uD2B8 \uC70C\uB9AC\uC5C4 \uD640\uB9AC(Robert William Holley, 1922\uB144 1\uC6D4 28\uC77C ~ 1993\uB144 2\uC6D4 11\uC77C)\uB294 \uBBF8\uAD6D\uC758 \uC0DD\uD654\uD559\uC790\uC774\uB2E4."@ko . . "Robert W. Holley (Urbana (Illinois), 28 de enero de 1922-Los Gatos (California), 11 de febrero de 1993) fue un bioqu\u00EDmico estadounidense, quien fue galardonado con el Premio Nobel de Fisiolog\u00EDa o Medicina en 1968, junto con Marshall Warren Nirenberg y Har Gobind Khorana por el descubrimiento de la estructura de alanina del ARN de transferencia, ligado al ADN y a la s\u00EDntesis proteica. En 1968 Holley se convirti\u00F3 en profesor residente del Instituto Salk por Estudios Biol\u00F3gicos en La Jolla (California)."@es . . . . . . "Bithcheimic\u00ED a rugadh in Urbana, Illinois ab ea Robert William Holley (28 Ean\u00E1ir 1922 \u2013 11 Feabhra 1993). Ar an bhfoireann a shint\u00E9isigh peinicillin don ch\u00E9ad uair sna 1940id\u00ED. I 1962 d'aithin s\u00E9 dh\u00E1 RNA traschuir don ch\u00E9ad uair, agus go luath ina dhiaidh sin d'aonraigh sampla de RNA traschuir \u00EDon don ch\u00E9ad uair. Sa bhliain 1965 d'fhoilsigh s\u00E9 strucht\u00FAr an aig\u00E9id n\u00FAicl\u00E9asaigh seo, at\u00E1 bun\u00FAsach i nd\u00E9anamh na bpr\u00F3it\u00E9in\u00ED sa chill. Le Har Gobind Khorana is Marshall Nirenberg, bhuaigh s\u00E9 Duais Nobel na Fiseola\u00EDochta n\u00F3 an Leighis sa bhliain 1968 as an saothar seo."@ga . "Robert William Holley (January 28, 1922 \u2013 February 11, 1993) was an American biochemist. He shared the Nobel Prize in Physiology or Medicine in 1968 (with Har Gobind Khorana and Marshall Warren Nirenberg) for describing the structure of alanine transfer RNA, linking DNA and protein synthesis. Using the Holley team's method, other scientists determined the structures of the remaining tRNA's. A few years later the method was modified to help track the sequence of nucleotides in various bacterial, plant, and human viruses. ."@en . . . "\u30ED\u30D0\u30FC\u30C8\u30FB\u30A6\u30A3\u30EA\u30A2\u30E0\u30FB\u30DB\u30EA\u30FC\uFF08Robert William Holley, 1922\u5E741\u670828\u65E5 - 1993\u5E742\u670811\u65E5\uFF09\u306F\u3001\u30A2\u30E1\u30EA\u30AB\u5408\u8846\u56FD\u306E\u751F\u5316\u5B66\u8005\u3067\u3001\u30A2\u30E9\u30CB\u30F3\u306E\u904B\u642CRNA\u3092\u4ECB\u3057\u3066DNA\u3068\u30BF\u30F3\u30D1\u30AF\u8CEA\u5408\u6210\u3092\u7D50\u3073\u3064\u3051\u305F\u7814\u7A76\u3067\u3001\u30CF\u30FC\u30FB\u30B4\u30D3\u30F3\u30C9\u30FB\u30B3\u30E9\u30CA\u3001\u30DE\u30FC\u30B7\u30E3\u30EB\u30FB\u30CB\u30FC\u30EC\u30F3\u30D0\u30FC\u30B0\u3068\u3068\u3082\u306B1968\u5E74\u5EA6\u306E\u30CE\u30FC\u30D9\u30EB\u751F\u7406\u5B66\u30FB\u533B\u5B66\u8CDE\u3092\u53D7\u8CDE\u3057\u305F\u3002"@ja . "Robert William Holley"@it . . . . . . . "Robert William Holley (* 28. Januar 1922 in Urbana, Illinois, USA; \u2020 11. Februar 1993 in Los Gatos, Kalifornien, USA) war ein US-amerikanischer Biochemiker und Nobelpreistr\u00E4ger. Holley war der Sohn von Lehrern. Er studierte ab 1938 Chemie an der University of Illinois at Urbana-Champaign mit dem Bachelor-Abschluss 1942. Er setzte sein Studium an der Cornell University fort mit der Dissertation zu einem Thema aus der organischen Chemie und der Promotion zum Ph. D. 1947 und war danach Dozent am State College of Washington in Seattle. 1948 wurde er Assistant Professor, 1957 Associate Professor und sp\u00E4ter Professor f\u00FCr Biochemie an der Cornell University, wobei er nebenbei bis 1964 als Forschungschemiker f\u00FCr das US Plant, Soil and Nutrition Laboratory arbeitete. Ab 1968 war er Resident Fellow am Salk Institute in La Jolla und daneben Adjunct Professor an der University of California, San Diego. Holley gelang im Jahr 1962 erstmals die Isolierung einer transfer-RNA, zwei Jahre sp\u00E4ter sequenzierte er die Alanin-tRNA. Das war die erste t-RNA, die sequenziert wurde (77 Nukleotide, sp\u00E4ter auf 76 korrigiert). Darauf aufbauend schlug er 1965 die Kleeblattstruktur der t-RNA vor, die sich auch bei anderen Vertretern der t-RNA best\u00E4tigte. Sp\u00E4ter arbeitete er \u00FCber die Kontrollsubstanzen der Zellteilung bei S\u00E4ugetieren. 1965 erhielt Holley den Albert Lasker Award for Basic Medical Research, 1966 wurde er in die American Academy of Arts and Sciences gew\u00E4hlt, 1968 in die National Academy of Sciences. 1967 erhielt er den National Academy of Sciences Award in Molecular Biology und 1968 gemeinsam mit Har Gobind Khorana und Marshall Warren Nirenberg den Nobelpreis f\u00FCr Physiologie oder Medizin f\u00FCr die Interpretation des genetischen Codes und seiner Funktion bei der Proteinsynthese."@de . . . "\u30ED\u30D0\u30FC\u30C8\u30FBW\u30FB\u30DB\u30EA\u30FC"@ja . . . . "Robert W. Holley"@en . . . . . . . . . . . . . . . "Robert W. Holley"@es . . "Robert William HOLLEY (28a de Januaro, 1922 \u2013 11a de Februaro, 1993) estis usona biokemiisto. Li kunhavis la Nobel-premion pri fiziologio a\u016D medicino en 1968 (kun Har Gobind Khorana kaj Marshall Warren Nirenberg) \u0109ar ili priskribis la strukturon de alanintransiga RNA, ligante DNA kaj proteina sintezo."@eo . "6692"^^ . . . . . . . . . "American"@en . "Robert William Holley"@in . . "Robert William Holley"@en . "\u0631\u0648\u0628\u0631\u062A \u0647\u0648\u0644\u064A"@ar . . . "\uB85C\uBC84\uD2B8 W. \uD640\uB9AC"@ko . "Robert William Holley (1922-1993) memerima Penghargaan Nobel dalam Fisiologi atau Kedokteran 1968 dengan Marshall Warren Nirenberg dan Hargobind Singh Khorana dan menjabat sebagai profesor di Universitas Cornell dan untuk kerjanya pada struktur ARN."@in . . . . . . . . . . . . . "Robert William Holley (* 28. Januar 1922 in Urbana, Illinois, USA; \u2020 11. Februar 1993 in Los Gatos, Kalifornien, USA) war ein US-amerikanischer Biochemiker und Nobelpreistr\u00E4ger. Holley war der Sohn von Lehrern. Er studierte ab 1938 Chemie an der University of Illinois at Urbana-Champaign mit dem Bachelor-Abschluss 1942. Er setzte sein Studium an der Cornell University fort mit der Dissertation zu einem Thema aus der organischen Chemie und der Promotion zum Ph. D. 1947 und war danach Dozent am State College of Washington in Seattle. 1948 wurde er Assistant Professor, 1957 Associate Professor und sp\u00E4ter Professor f\u00FCr Biochemie an der Cornell University, wobei er nebenbei bis 1964 als Forschungschemiker f\u00FCr das US Plant, Soil and Nutrition Laboratory arbeitete. Ab 1968 war er Resident Fellow "@de . . . . "Robert W. Holley"@sv . . . . "Robert William Holley (Urbana, 28 de janeiro de 1922 \u2014 Los Gatos, 11 de fevereiro de 1993) foi um bioqu\u00EDmico estadunidense. Compartilhou o Pr\u00EAmio Nobel de Fisiologia ou Medicina em 1968 (com Har Gobind Khorana e Marshall Nirenberg) por descrever a estrutura do RNA de transfer\u00EAncia de alanina, ligando o DNA e a s\u00EDntese de prote\u00EDnas. Holley nasceu em Urbana, Illinois, e se formou na Urbana High School em 1938. Ele passou a estudar qu\u00EDmica na Universidade de Illinois em Urbana-Champaign, graduando-se em 1942 e iniciando seus estudos de PhD em qu\u00EDmica org\u00E2nica na Cornell University. Durante a Segunda Guerra Mundial, Holley passou dois anos trabalhando com o professor Vincent du Vigneaud no Cornell University Medical College, onde esteve envolvido na primeira s\u00EDntese qu\u00EDmica da penicilina. Holley concluiu seus estudos de PhD em 1947. Ap\u00F3s seus estudos de gradua\u00E7\u00E3o, Holley continuou associado a Cornell. Ele se tornou professor assistente de qu\u00EDmica org\u00E2nica em 1948 e foi nomeado professor de bioqu\u00EDmica em 1962. Ele come\u00E7ou sua pesquisa sobre RNA depois de passar um ano sab\u00E1tico (1955-1956) estudando com James F. Bonner no Instituto de Tecnologia da Calif\u00F3rnia. A pesquisa de Holley sobre o RNA se concentrou primeiro no isolamento do RNA de transfer\u00EAncia (tRNA) e, posteriormente, na determina\u00E7\u00E3o da sequ\u00EAncia e estrutura do tRNA da alanina, a mol\u00E9cula que incorpora o amino\u00E1cido alanina nas prote\u00EDnas. A equipe de pesquisadores de Holley determinou a estrutura do tRNA usando duas ribonucleases para dividir a mol\u00E9cula de tRNA em peda\u00E7os. Cada enzima divide a mol\u00E9cula em pontos de localiza\u00E7\u00E3o para nucleot\u00EDdeos espec\u00EDficos. Por um processo de \"confundir\" a estrutura das pe\u00E7as divididas pelas duas enzimas diferentes e, em seguida, comparando as pe\u00E7as de ambas as divis\u00F5es de enzimas, a equipe acabou determinando toda a estrutura da mol\u00E9cula. O grupo de pesquisadores inclui Elizabeth Beach Keller, que desenvolveu o modelo em folha de trevo que descreve o RNA de transfer\u00EAncia, durante o curso da pesquisa. A estrutura foi conclu\u00EDda em 1964, e foi uma descoberta importante para explicar a s\u00EDntese de prote\u00EDnas a partir do RNA mensageiro. Foi tamb\u00E9m a primeira sequ\u00EAncia de nucleot\u00EDdeos de um \u00E1cido ribonucleico j\u00E1 determinada. Holley recebeu o Pr\u00EAmio Nobel de Fisiologia ou Medicina em 1968 por esta descoberta, e Har Gobind Khorana e Marshall W. Nirenberg tamb\u00E9m receberam o pr\u00EAmio naquele ano por contribui\u00E7\u00F5es para a compreens\u00E3o da s\u00EDntese de prote\u00EDnas. Usando o m\u00E9todo da equipe de Holley, outros cientistas determinaram as estruturas dos tRNAs restantes. Alguns anos mais tarde, o m\u00E9todo foi modificado para ajudar a rastrear a sequ\u00EAncia de nucleot\u00EDdeos em v\u00E1rios v\u00EDrus bacterianos, vegetais e humanos. Em 1968, Holley tornou-se um membro residente do Instituto Salk de Estudos Biol\u00F3gicos em La Jolla, Calif\u00F3rnia."@pt . . . "1096424619"^^ . "Robert William Holley (January 28, 1922 \u2013 February 11, 1993) was an American biochemist. He shared the Nobel Prize in Physiology or Medicine in 1968 (with Har Gobind Khorana and Marshall Warren Nirenberg) for describing the structure of alanine transfer RNA, linking DNA and protein synthesis. Holley was born in Urbana, Illinois, and graduated from Urbana High School in 1938. He went on to study chemistry at the University of Illinois at Urbana-Champaign, graduating in 1942 and commencing his PhD studies in organic chemistry at Cornell University. During World War II Holley spent two years working under Professor Vincent du Vigneaud at Cornell University Medical College, where he was involved in the first chemical synthesis of penicillin. Holley completed his PhD studies in 1947. Following his graduate studies Holley remained associated with Cornell. He became an assistant professor of organic chemistry in 1948, and was appointed as professor of biochemistry in 1962. He began his research on RNA after spending a year's sabbatical (1955\u20131956) studying with James F. Bonner at the California Institute of Technology. Holley's research on RNA focused first on isolating transfer RNA (tRNA), and later on determining the sequence and structure of alanine tRNA, the molecule that incorporates the amino acid alanine into proteins. Holley's team of researchers determined the tRNA's structure by using two ribonucleases to split the tRNA molecule into pieces. Each enzyme split the molecule at location points for specific nucleotides. By a process of \"puzzling out\" the structure of the pieces split by the two different enzymes, then comparing the pieces from both enzyme splits, the team eventually determined the entire structure of the molecule. The group of researchers include Elizabeth Beach Keller, who developed the cloverleaf model that describes transfer RNA, during the course of the research. The structure was completed in 1964, and was a key discovery in explaining the synthesis of proteins from messenger RNA. It was also the first nucleotide sequence of a ribonucleic acid ever determined. Holley was awarded the Nobel Prize in Physiology or Medicine in 1968 for this discovery, and Har Gobind Khorana and Marshall W. Nirenberg were also awarded the prize that year for contributions to the understanding of protein synthesis. Using the Holley team's method, other scientists determined the structures of the remaining tRNA's. A few years later the method was modified to help track the sequence of nucleotides in various bacterial, plant, and human viruses. In 1968 Holley became a resident fellow at the Salk Institute for Biological Studies in La Jolla, California. According to the New York Times obituary, \"He was an avid outdoorsman and an amateur sculptor of bronze.\" ."@en . . "Robert W. Holley"@eo . . "Robert William Holley (Urbana, Illinois, EUA 1922 - Los Gatos 1993) fou un bioqu\u00EDmic i professor universitari nord-americ\u00E0 guardonat amb el Premi Nobel de Medicina o Fisiologia l'any 1968."@ca . . "Robert W. Holley"@nl . "1922-01-28"^^ . . . . "Robert W. Holley"@ca . . . . . "Robert William Holley (Urbana, Illinois, AEB, 1922ko urtarrilaren 28a - Los Gatos, Kalifornia, 1993ko otsailaren 11) Estatu Batuetako biokimikaria izan zen."@eu . . . . . . . "\u30ED\u30D0\u30FC\u30C8\u30FB\u30A6\u30A3\u30EA\u30A2\u30E0\u30FB\u30DB\u30EA\u30FC\uFF08Robert William Holley, 1922\u5E741\u670828\u65E5 - 1993\u5E742\u670811\u65E5\uFF09\u306F\u3001\u30A2\u30E1\u30EA\u30AB\u5408\u8846\u56FD\u306E\u751F\u5316\u5B66\u8005\u3067\u3001\u30A2\u30E9\u30CB\u30F3\u306E\u904B\u642CRNA\u3092\u4ECB\u3057\u3066DNA\u3068\u30BF\u30F3\u30D1\u30AF\u8CEA\u5408\u6210\u3092\u7D50\u3073\u3064\u3051\u305F\u7814\u7A76\u3067\u3001\u30CF\u30FC\u30FB\u30B4\u30D3\u30F3\u30C9\u30FB\u30B3\u30E9\u30CA\u3001\u30DE\u30FC\u30B7\u30E3\u30EB\u30FB\u30CB\u30FC\u30EC\u30F3\u30D0\u30FC\u30B0\u3068\u3068\u3082\u306B1968\u5E74\u5EA6\u306E\u30CE\u30FC\u30D9\u30EB\u751F\u7406\u5B66\u30FB\u533B\u5B66\u8CDE\u3092\u53D7\u8CDE\u3057\u305F\u3002"@ja . . . . . . . . . . "\u0420\u043E\u0301\u0431\u0435\u0440\u0442 \u0423\u0438\u0301\u043B\u044C\u044F\u043C \u0425\u043E\u0301\u043B\u043B\u0438 (\u0430\u043D\u0433\u043B. Robert William Holley; 28 \u044F\u043D\u0432\u0430\u0440\u044F 1922, \u042D\u0440\u0431\u0430\u043D\u0430, \u0418\u043B\u043B\u0438\u043D\u043E\u0439\u0441 \u2014 11 \u0444\u0435\u0432\u0440\u0430\u043B\u044F 1993, \u041B\u043E\u0441-\u0413\u0430\u0442\u043E\u0441, \u041A\u0430\u043B\u0438\u0444\u043E\u0440\u043D\u0438\u044F) \u2014 \u0430\u043C\u0435\u0440\u0438\u043A\u0430\u043D\u0441\u043A\u0438\u0439 \u0431\u0438\u043E\u0445\u0438\u043C\u0438\u043A, \u043B\u0430\u0443\u0440\u0435\u0430\u0442 \u041D\u043E\u0431\u0435\u043B\u0435\u0432\u0441\u043A\u043E\u0439 \u043F\u0440\u0435\u043C\u0438\u0438 \u043F\u043E \u0444\u0438\u0437\u0438\u043E\u043B\u043E\u0433\u0438\u0438 \u0438 \u043C\u0435\u0434\u0438\u0446\u0438\u043D\u0435 \u0432 1968 \u0433\u043E\u0434\u0443 (\u0441\u043E\u0432\u043C\u0435\u0441\u0442\u043D\u043E \u0441 \u0425\u0430\u0440\u043E\u043C \u041A\u043E\u0440\u0430\u043D\u0430 \u0438 \u041C\u0430\u0440\u0448\u0430\u043B\u043B\u043E\u043C \u041D\u0438\u0440\u0435\u043D\u0431\u0435\u0440\u0433\u043E\u043C) \u00AB\u0437\u0430 \u0440\u0430\u0441\u0448\u0438\u0444\u0440\u043E\u0432\u043A\u0443 \u0433\u0435\u043D\u0435\u0442\u0438\u0447\u0435\u0441\u043A\u043E\u0433\u043E \u043A\u043E\u0434\u0430 \u0438 \u0435\u0433\u043E \u0440\u043E\u043B\u0438 \u0432 \u0441\u0438\u043D\u0442\u0435\u0437\u0435 \u0431\u0435\u043B\u043A\u043E\u0432\u00BB. \u0427\u043B\u0435\u043D \u041D\u0430\u0446\u0438\u043E\u043D\u0430\u043B\u044C\u043D\u043E\u0439 \u0430\u043A\u0430\u0434\u0435\u043C\u0438\u0438 \u043D\u0430\u0443\u043A \u0421\u0428\u0410 (1968)."@ru . "\u7F85\u4F2F\u7279\u00B7\u5A01\u5EC9\u00B7\u970D\u5229\uFF08\u82F1\u8A9E\uFF1ADr Robert William Holley\uFF0C1922\u5E741\u670828\u65E5\uFF0D1993\u5E742\u670811\u65E5\uFF09\uFF0C\u7F8E\u570B\u751F\u7269\u5316\u5B78\u5BB6\uFF0C\u51FA\u751F\u65BC\u4F0A\u5229\u8AFE\u5DDE\u5384\u5DF4\u7D0D\u3002\u66FE\u7372\u5F971968\u5E74\u8AFE\u8C9D\u723E\u751F\u7406\u5B78\u6216\u91AB\u5B78\u734E\uFF0C\u539F\u56E0\u662F\u95E1\u660E\u4E86\u9023\u7D50\u4E19\u6C28\u9178\u8207DNA\u7684tRNA\u3002\u5171\u540C\u7372\u734E\u4EBA\u9084\u6709\u54C8\u723E\u00B7\u845B\u8CD3\u00B7\u79D1\u62C9\u7D0D\uFF08Har Gobind Khorana\uFF09\u53CA\u99AC\u6B47\u723E\u00B7\u6C83\u502B\u00B7\u5C3C\u502B\u4F2F\u683C\uFF08Marshall Warren Nirenberg\uFF09\u3002"@zh . . . "Robert William Holley, n\u00E9 le 28 janvier 1922 \u00E0 Urbana dans l'Illinois aux \u00C9tats-Unis et d\u00E9c\u00E9d\u00E9 le 11 f\u00E9vrier 1993, est un chimiste organique et biochimiste am\u00E9ricain. Il re\u00E7oit le prix Nobel de m\u00E9decine en 1968."@fr . . "Bithcheimic\u00ED a rugadh in Urbana, Illinois ab ea Robert William Holley (28 Ean\u00E1ir 1922 \u2013 11 Feabhra 1993). Ar an bhfoireann a shint\u00E9isigh peinicillin don ch\u00E9ad uair sna 1940id\u00ED. I 1962 d'aithin s\u00E9 dh\u00E1 RNA traschuir don ch\u00E9ad uair, agus go luath ina dhiaidh sin d'aonraigh sampla de RNA traschuir \u00EDon don ch\u00E9ad uair. Sa bhliain 1965 d'fhoilsigh s\u00E9 strucht\u00FAr an aig\u00E9id n\u00FAicl\u00E9asaigh seo, at\u00E1 bun\u00FAsach i nd\u00E9anamh na bpr\u00F3it\u00E9in\u00ED sa chill. Le Har Gobind Khorana is Marshall Nirenberg, bhuaigh s\u00E9 Duais Nobel na Fiseola\u00EDochta n\u00F3 an Leighis sa bhliain 1968 as an saothar seo."@ga . . . . "Robert William Holley, n\u00E9 le 28 janvier 1922 \u00E0 Urbana dans l'Illinois aux \u00C9tats-Unis et d\u00E9c\u00E9d\u00E9 le 11 f\u00E9vrier 1993, est un chimiste organique et biochimiste am\u00E9ricain. Il re\u00E7oit le prix Nobel de m\u00E9decine en 1968."@fr . "Robert Holley"@fr . . . . "Robert William Holley (Urbana (Illinois) 28 januari 1922 \u2013 Los Gatos 11 februari 1993) was een Amerikaans biochemicus en Nobelprijswinnaar. In 1968 won hij samen met Har Gobind Khorana en Marshall Nirenberg de Nobelprijs voor de Fysiologie of Geneeskunde voor het beschrijven van de structuur van alanine transfer RNA, verbindend DNA en prote\u00EFnesynthese."@nl . . . "Robert William Holley (1922-1993) memerima Penghargaan Nobel dalam Fisiologi atau Kedokteran 1968 dengan Marshall Warren Nirenberg dan Hargobind Singh Khorana dan menjabat sebagai profesor di Universitas Cornell dan untuk kerjanya pada struktur ARN. Holley lulus dari Universitas Illinois pada 1942 dan meneruskan menerima PhD dalam kimia organik dari Cornell University pada 1947. Ia bekerja di sana sebagai guru besar dan kimiawan peneliti di dan hingga 1966, saat ia pindah ke dan di . Ia bergabung dengan fakultas di Salk Institute sebagai anggota residen dan profesor biologi molekuler pada 1968. Holley menerima beberapa penghargaan selain Hadiah Nobel, termasuk yang prestisius pada 1965."@in . . . . "Robert W. Holley"@en . "Robert William Holley (Urbana, 28 gennaio 1922 \u2013 Los Gatos, 11 febbraio 1993) \u00E8 stato un biochimico statunitense, noto per aver individuato la struttura del tRNA responsabile del trasporto dell'alanina ed aver contribuito, insieme a insieme a Marshall Nirenberg ed Har Gobind Khorana, a descrivere il meccanismo della sintesi proteica. Per tali ricerche, insieme a Nirenberg e Khorana, Holley vinse il Premio Nobel per la Medicina nel 1968."@it . . . "\u0420\u043E\u0431\u0435\u0440\u0442 \u0412\u0456\u043B\u044C\u044F\u043C \u0413\u043E\u043B\u043B\u0456"@uk . "1993-02-11"^^ . . "Robert W. Holley (Urbana (Illinois), 28 de enero de 1922-Los Gatos (California), 11 de febrero de 1993) fue un bioqu\u00EDmico estadounidense, quien fue galardonado con el Premio Nobel de Fisiolog\u00EDa o Medicina en 1968, junto con Marshall Warren Nirenberg y Har Gobind Khorana por el descubrimiento de la estructura de alanina del ARN de transferencia, ligado al ADN y a la s\u00EDntesis proteica. Holley se gradu\u00F3 de la secundaria de su ciudad natal en 1938. Realiz\u00F3 sus estudios en Qu\u00EDmica en el campus de la Universidad de Illinois at Urbana-Champaign, gradu\u00E1ndose en 1942 y comenzando sus estudios de doctorado en Qu\u00EDmica Org\u00E1nica en la Universidad de Cornell. Durante la Segunda Guerra Mundial, Holley pas\u00F3 dos a\u00F1os trabajando bajo la supervisi\u00F3n del profesor Vincent du Vigneaud en la Unidad M\u00E9dica de la Universidad de Cornell, donde fue part\u00EDcipe de la primera s\u00EDntesis qu\u00EDmica de la penicilina. Complet\u00F3 sus estudios de doctorado en 1947. En 1968 Holley se convirti\u00F3 en profesor residente del Instituto Salk por Estudios Biol\u00F3gicos en La Jolla (California)."@es . "Robert William Holley (Urbana, Illinois, AEB, 1922ko urtarrilaren 28a - Los Gatos, Kalifornia, 1993ko otsailaren 11) Estatu Batuetako biokimikaria izan zen."@eu . "Robert William Holley (Urbana, 28 gennaio 1922 \u2013 Los Gatos, 11 febbraio 1993) \u00E8 stato un biochimico statunitense, noto per aver individuato la struttura del tRNA responsabile del trasporto dell'alanina ed aver contribuito, insieme a insieme a Marshall Nirenberg ed Har Gobind Khorana, a descrivere il meccanismo della sintesi proteica. Per tali ricerche, insieme a Nirenberg e Khorana, Holley vinse il Premio Nobel per la Medicina nel 1968."@it . . . . . . . . . . . . "\u0420\u043E\u0301\u0431\u0435\u0440\u0442 \u0412\u0456\u0301\u043B\u044C\u044F\u043C \u0413\u043E\u0301\u043B\u043B\u0456 (\u0430\u043D\u0433\u043B. Robert William Holley; 28 \u0441\u0456\u0447\u043D\u044F 1922, \u0423\u0440\u0431\u0430\u043D\u0430, \u0406\u043B\u043B\u0456\u043D\u043E\u0439\u0441 \u2014 11 \u043B\u044E\u0442\u043E\u0433\u043E 1993, \u041B\u043E\u0441-\u0413\u0430\u0442\u043E\u0441, \u041A\u0430\u043B\u0456\u0444\u043E\u0440\u043D\u0456\u044F, \u0421\u0428\u0410) \u2014 \u0430\u043C\u0435\u0440\u0438\u043A\u0430\u043D\u0441\u044C\u043A\u0438\u0439 \u0431\u0456\u043E\u0445\u0456\u043C\u0456\u043A, \u043B\u0430\u0432\u0440\u0435\u0430\u0442 \u041D\u043E\u0431\u0435\u043B\u0456\u0432\u0441\u044C\u043A\u043E\u0457 \u043F\u0440\u0435\u043C\u0456\u0457 \u0437 \u043C\u0435\u0434\u0438\u0446\u0438\u043D\u0438 \u0442\u0430 \u0444\u0456\u0437\u0456\u043E\u043B\u043E\u0433\u0456\u0457 1968 \u0440\u043E\u043A\u0443 (\u0441\u043F\u0456\u043B\u044C\u043D\u043E \u0437 \u0413\u0430\u0440\u043E\u043C \u0425\u043E\u0440\u0430\u043D\u0430 \u0456 \u041C\u0430\u0440\u0448\u0430\u043B\u043B\u043E\u043C \u041D\u0456\u0440\u0435\u043D\u0431\u0435\u0440\u0433\u043E\u043C) \u00AB\u0437\u0430 \u0440\u043E\u0437\u0448\u0438\u0444\u0440\u043E\u0432\u043A\u0443 \u0433\u0435\u043D\u0435\u0442\u0438\u0447\u043D\u043E\u0433\u043E \u043A\u043E\u0434\u0443 \u0456 \u0439\u043E\u0433\u043E \u0440\u043E\u043B\u0456 \u0432 \u0441\u0438\u043D\u0442\u0435\u0437\u0456 \u0431\u0456\u043B\u043A\u0456\u0432\u00BB, \u0437\u043E\u043A\u0440\u0435\u043C\u0430 \u0432\u043D\u0435\u0441\u043A\u043E\u043C \u0413\u043E\u043B\u043B\u0456 \u0431\u0443\u0432 \u043E\u043F\u0438\u0441 \u043F\u043E\u0441\u043B\u0456\u0434\u043E\u0432\u043D\u043E\u0441\u0442\u0456 \u0439 \u0441\u0442\u0440\u0443\u043A\u0442\u0443\u0440\u0438 \u0442\u0420\u041D\u041A \u0430\u043B\u0430\u043D\u0456\u043D\u0443, \u043F\u0435\u0440\u0448\u043E\u0457 \u043C\u043E\u043B\u0435\u043A\u0443\u043B\u0438 \u0420\u041D\u041A, \u0434\u043B\u044F \u044F\u043A\u043E\u0457 \u0431\u0443\u043B\u0430 \u0432\u0441\u0442\u0430\u043D\u043E\u0432\u043B\u0435\u043D\u0430 \u0441\u0442\u0440\u0443\u043A\u0442\u0443\u0440\u0430."@uk . . "\u0631\u0648\u0628\u0631\u062A \u0648\u0644\u064A\u0627\u0645 \u0647\u0648\u0644\u064A (\u0628\u0627\u0644\u0625\u0646\u062C\u0644\u064A\u0632\u064A\u0629: Robert W. Holley)\u200F \u0643\u0627\u0646 \u0639\u0627\u0644\u0645 \u0643\u064A\u0645\u064A\u0627\u0621 \u062D\u064A\u0648\u064A\u0629 \u0623\u0645\u0631\u064A\u0643\u064A (28 \u064A\u0646\u0627\u064A\u0631 1922 - 11 \u0641\u0628\u0631\u0627\u064A\u0631 1993)\u060C \u062A\u062D\u0635\u0644 \u0639\u0644\u0649 \u062C\u0627\u0626\u0632\u0629 \u0646\u0648\u0628\u0644 \u0641\u064A \u0627\u0644\u0637\u0628 \u0644\u0639\u0627\u0645 1968 \u0645\u0634\u0627\u0631\u0643\u0629 \u0645\u0639 \u0647\u0627\u0631 \u063A\u0648\u0628\u0646\u062F \u062E\u0648\u0631\u0627\u0646\u0627 \u0648\u0645\u0627\u0631\u0634\u0627\u0644 \u0646\u064A\u0631\u0646\u0628\u0631\u063A \u0648\u0630\u0644\u0643 \u0644\u0623\u0628\u062D\u0627\u062B\u0647\u0645 \u0627\u0644\u062A\u064A \u0623\u0644\u0642\u062A \u0627\u0644\u0636\u0648\u0621 \u0639\u0644\u0649 \u0627\u0644\u0645\u0643\u0648\u0646\u0627\u062A \u0627\u0644\u0648\u0631\u0627\u062B\u064A\u0629 \u0627\u0644\u0645\u0633\u0626\u0648\u0644\u0629 \u0639\u0646 \u062A\u0646\u0638\u064A\u0645 \u0639\u0645\u0644\u064A\u0629 \u062A\u062E\u0644\u064A\u0642 \u0627\u0644\u0628\u0631\u0648\u062A\u064A\u0646\u0627\u062A \u0641\u064A \u0646\u0648\u0627\u0629 \u0627\u0644\u062E\u0644\u064A\u0629."@ar . . . . "Robert William Holley (ur. 28 stycznia 1922 w Urbana, zm. 11 lutego 1993 w Los Gatos) \u2013 ameryka\u0144ski biochemik, laureat Nagrody Nobla z dziedziny fizjologii lub medycyny w roku 1968 za wyja\u015Bnienie istoty kodu genetycznego i ustalenie struktury tRNA. W latach 1948-1969 profesor Cornell University w Ithaca i w La Jolla (San Diego). Prowadzi\u0142 badania syntezy bia\u0142ek i kwas\u00F3w nukleinowych. Za ustalenie struktury kwasu rybonukleinowego przenosz\u0105cego tRNA i niezale\u017Cne wyja\u015Bnienie istoty kodu genetycznego oraz jego roli w biosyntezie bia\u0142ka otrzyma\u0142 w 1968 roku Nagrod\u0119 Nobla (razem z Harem Khoran\u0105 i Marshallem W. Nirenbergiem)."@pl . . "Robert Holley"@ga . . . . . . . . "Robert William Holley (28. ledna 1922 \u2013 11. \u00FAnora 1993) byl americk\u00FD biochemik a chemik, nositel Nobelovy ceny za fyziologii a l\u00E9ka\u0159stv\u00ED za rok 1968. Spolu s n\u00EDm ji z\u00EDskali Har Gobind Khorana a Marshall Warren Nirenberg a cena byla ud\u011Blena za vysv\u011Btlen\u00ED toho, jak sekvence nukleotid\u016F v nukleov\u00FDch kyselin\u00E1ch \u0159\u00EDd\u00ED synt\u00E9zu protein\u016F v bu\u0148k\u00E1ch. V\u011Bt\u0161inu sv\u00E9 v\u011Bdeck\u00E9 kari\u00E9ry str\u00E1vil Robert W. Holley na Cornellov\u011B univerzit\u011B, kde i studoval."@cs . "Robert William Holley (Urbana, 28 de janeiro de 1922 \u2014 Los Gatos, 11 de fevereiro de 1993) foi um bioqu\u00EDmico estadunidense. Compartilhou o Pr\u00EAmio Nobel de Fisiologia ou Medicina em 1968 (com Har Gobind Khorana e Marshall Nirenberg) por descrever a estrutura do RNA de transfer\u00EAncia de alanina, ligando o DNA e a s\u00EDntese de prote\u00EDnas. Usando o m\u00E9todo da equipe de Holley, outros cientistas determinaram as estruturas dos tRNAs restantes. Alguns anos mais tarde, o m\u00E9todo foi modificado para ajudar a rastrear a sequ\u00EAncia de nucleot\u00EDdeos em v\u00E1rios v\u00EDrus bacterianos, vegetais e humanos."@pt . . "Robert William Holley"@en . . "\u0425\u043E\u043B\u043B\u0438, \u0420\u043E\u0431\u0435\u0440\u0442"@ru . . . . . . . . . . . . . . . . . "Robert William Holley"@eu . . . "\u7F85\u4F2F\u7279\u00B7W\u00B7\u970D\u5229"@zh . "\uB85C\uBC84\uD2B8 \uC70C\uB9AC\uC5C4 \uD640\uB9AC(Robert William Holley, 1922\uB144 1\uC6D4 28\uC77C ~ 1993\uB144 2\uC6D4 11\uC77C)\uB294 \uBBF8\uAD6D\uC758 \uC0DD\uD654\uD559\uC790\uC774\uB2E4."@ko . .