Ruth Clayton (née Freedman;10 June 1925 – 2003) was a lecturer and researcher with an international reputation in eye research at the University of Edinburgh in the Institute of Animal Genetics. She was one of a group of scientists who joined the Institute at a time when it was led by C.H. Waddington and when many of the fundamental aspects of modern biology were being elucidated by forward thinking scientists at the university's King's Buildings campus. Leading a large and diverse research group, she applied the newly developing techniques of modern biology to fundamental questions in the fields of developmental biology and pathology of the eye and the brain. Her work was characterised by a rigorously conceptual approach, methodological innovation and a keen interest in the social and eth
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